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Melanocyte-stimulating hormone is a derivative of the precursor proopiomelanocortin, an agonist on the melanocortin-1 receptor that promotes formation of eumelanin

increase, decrease, tumor inflammatory genes Gremlin (GREM1), interleukin 1 (IL-1), interleukin-4 [IL-4], nuclear factor B subunit 1 (NFB1), and prostaglandin-endoperoxide synthase 2 (PTGS2), extracellular signal-regulated kinase (ERK), c-jun N-terminal kinase (JNK), light chain 3 protein B II (LC3BII), c-Jun N-terminal kinase (JNK), poly(ADP ribose) polymerase (PARP)-1, nuclear factor-kappa B (NF-kappaB), cyclooxygenase-2 (Cox-2), extracellular signal-regulated protein kinase (ERK1/2), mitogen-activated protein kinase (MAPK), 7,12-dimethylbenz(a)anthracene (DMBA)-initiated and 12-O-tetradecanoylphorbol-13-acetate (TPA), metalloproteinases MMP